Angiosperms (flowering plants) are the largest group in the plant kingdom, and they are broadly classified into two categories: monocots and dicots. In this article, we will focus on the anatomy of dicotyledonous and monocotyledonous plants, with special emphasis on the structure of dicot and monocot root, stem and leaf. We will also explore differences and key characteristics and provide a diagram overview to help you understand both types more clearly.
Root System
Dicots usually have a taproot system, which is a main root growing deep into the soil with lateral branches.
Epidermis
The outermost layer, called the epidermis, sometimes bears root hairs for water and mineral absorption.
Cortex
Below the epidermis lies the cortex, composed of loosely packed parenchyma cells with intercellular spaces. This region helps in the storage and transport of water and minerals.
Endodermis
The innermost layer of the cortex is characterised by barrel-shaped, tightly packed cells. It regulates the flow of substances into the vascular cylinder.
Pericycle
A few layers of thick-walled parenchyma lie immediately inside the endodermis. In dicots, the pericycle is the origin of lateral roots and contributes to secondary growth.
Vascular Bundle
In most dicot roots, there are two to four xylem and phloem strands arranged in a radial pattern.
The xylem and phloem are separated by parenchymatous conjunctive tissue. During secondary growth, a cambium develops between the xylem and phloem, leading to an increase in root girth.
Pith
In many dicot roots, the pith at the centre may be small or inconspicuous.
Root System
Monocots typically have an adventitious root system. Many thin roots emerge from the stem base or nodes close to the soil surface.
Epidermis and Cortex
The structure of dicot and monocot root, stem and leaf can appear quite similar in the outermost layers. In monocot roots, the epidermis is followed by a multilayered cortex of parenchyma cells.
Endodermis and Pericycle
The endodermis is well-defined, followed by a pericycle. However, secondary growth is generally absent in monocot roots, so the pericycle does not form a vascular cambium as in dicots.
Vascular Bundles
In monocots, the number of xylem strands is usually six or more (polyarch condition). Xylem and phloem alternate with each other in a ring-like arrangement.
Pith
Monocots often have a large and distinct pith in the centre of the root.
No Secondary Growth
Unlike dicots, monocots do not exhibit secondary growth in their roots.
A dicot stem is usually solid and exhibits distinct regions when viewed in cross-section:
Epidermis
The outermost protective layer is often covered by a thin cuticle. Epidermal cells may bear multicellular hairs (trichomes) and a few stomata.
Cortex
The cortex is typically divided into three sub-zones:
Hypodermis: Often made of collenchymatous cells, providing mechanical support.
Middle Cortical Layers: Composed of parenchyma with intercellular spaces for storage and transport.
Endodermis: The innermost boundary of the cortex. Also called the starch sheath in many dicots.
Pericycle
Situated beneath the endodermis. In stems, it may include patches of sclerenchyma.
Vascular Bundles
In a typical young dicot stem, vascular bundles are arranged in a ring. This arrangement is often referred to as a ring or circular pattern.
Each vascular bundle is conjoint (xylem and phloem together), open (a cambium present between xylem and phloem), and has an endarch protoxylem (protoxylem towards the centre).
This cambium is responsible for secondary growth, leading to the formation of annual rings.
Pith
The central region of the stem is composed of parenchyma cells that store nutrients and water.
A monocot stem is usually hollow or fibrous, and it does not undergo significant secondary growth:
Epidermis
A single-layered covering with a thick cuticle.
Hypodermis
Often consists of sclerenchyma, making the stem firm.
Vascular Bundles
The vascular bundles are numerous and scattered throughout the ground tissue (which includes both cortex and pith together). This scattered arrangement is a key feature in monocot vs dicot comparisons.
Vascular bundles are conjoint (xylem and phloem together) and closed (no cambium present), so there is no secondary growth.
Phloem parenchyma is generally absent.
Ground Tissue
Unlike dicots, monocots often lack a well-demarcated cortex and pith. Instead, they have a uniform ground tissue made up of parenchymatous cells.
Studying the anatomy of the monocot stem is crucial to understanding how monocots support themselves without secondary thickening.
Dicot leaves typically show reticulate (net-like) venation and can be divided into the following layers:
Epidermis
The upper (adaxial) and lower (abaxial) epidermal layers are usually covered by a cuticle.
Stomata may be present on both surfaces, though they are generally more abundant on the lower surface. Some dicot leaves may have stomata only on the lower surface.
Mesophyll
Consists of palisade parenchyma (elongated cells rich in chloroplasts, arranged just under the upper epidermis) and spongy parenchyma (loosely arranged cells with air spaces near the lower epidermis).
The mesophyll is the primary site of photosynthesis.
Vascular Bundles
Form the veins and midrib of the leaf, surrounded by a bundle sheath of parenchyma cells.
The vein arrangement (reticulate venation) distinguishes it from parallel venation in monocots.
Monocot leaves are characterised by parallel venation:
Epidermis
Both upper (adaxial) and lower (abaxial) epidermal layers bear stomata, often arranged in rows.
Mesophyll
Typically, it is not differentiated into palisade and spongy parenchyma. The mesophyll is homogeneous.
Bulliform Cells
Large, thin-walled cells on the adaxial side help in leaf folding or rolling to reduce water loss during stress.
Vascular Bundles
Arranged parallel to each other, reflecting the parallel venation pattern.
By comparing the structure of dicot and monocot root, stem and leaf, you can see how these two groups differ in external morphology and internal arrangement of tissues.
Below is a concise overview of monocot vs dicot differences for roots, stems, and leaves:
Root: Taproot in dicots vs adventitious roots in monocots.
Stem: Vascular bundles in a ring (dicots) vs scattered bundles (monocots).
Leaf: Reticulate venation (dicots) vs parallel venation (monocots).
Test your knowledge of the anatomy of dicotyledonous and monocotyledonous plants with these short questions. Answers are provided at the end.
1. Which layer in the roots is responsible for regulating the entry of water and minerals into the vascular bundle?
a) Cortex
b) Endodermis
c) Pericycle
d) Epidermis
2. What is the typical arrangement of vascular bundles in a dicot stem?
a) Scattered in the ground tissue
b) Concentric circles without cambium
c) Ring arrangement with cambium
d) Randomly placed with phloem parenchyma
3. In which type of leaf (dicot or monocot) are bulliform cells found?
a) Dicot leaves
b) Monocot leaves
4. Which type of stem usually has open vascular bundles?
a) Dicot stem
b) Monocot stem
b) Endodermis
c) Ring arrangement with cambium
b) Monocot leaves
a) Dicot stem
1: Why is there no secondary growth in monocot roots and stems?
Monocots have closed vascular bundles without a vascular cambium layer. Since cambium is responsible for producing new xylem and phloem, monocots generally do not exhibit secondary growth.
2: How many xylem bundles are commonly found in monocot roots?
Monocot roots often possess six or more xylem bundles, a condition referred to as polyarch, which distinguishes them from dicots.
3: Why is the dicot vascular bundle called ‘open’?
In dicot stems, vascular bundles have a cambium between the xylem and phloem. This cambium remains active and adds a new xylem and phloem, making the bundle ‘open’ to secondary growth.
4: Do bulliform cells exist in dicot leaves?
No, bulliform cells are typically found only in certain monocot leaves. They help in rolling or folding the leaf to minimise water loss.
5: What is the main difference in venation patterns of monocot and dicot leaves?
Monocot leaves show parallel venation (veins run parallel), while dicot leaves have reticulate venation (veins form a network).